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period 3 = Feb to Mar; see table A.7 in Annexes). Cormack-Jolly-Seber (CJS) models are based on live captures and releases of marked individuals into the population. We therefore analysed the effects of cohort (1 and 2), age at first capture (juvenile and adult) and period on the return ratę by testing for an effect of group (group 1 = cohort 1/Adult, group 2 = cohort 1/Juvenile; group 3 = cohort 2/Adult; group 4 = cohort 2/ Juvenile) and period on both survival and encounter probabilities (return ratę = 0(grt)P(gict))-
For among year survival analyses (n = 56), we only used data firom cohort 1. We grouped encounter data in seven periods (period 1 = Sept to Nov 2009, period 2 = Dec 2009 to Jan 2010; period 3 = Feb to Mar 2010, period 4 = Sept to Nov 2010, period 5 = Dec 2010 to Jan 2011; period 6 = Feb to Mar 2011, period 7 = Nov to Dec 2011; see table A.8 in Annexes). We analyzed the relationship between age at first capture, periods and return ratę by testing the effect of group (group 1 = Adult; group 2 = Juvenile) and period on both survival and encounter probabilities (return ratę = 0(gx,)P(gXt)).
We first tested whether our data fitted the fuli time-dependent CJS model (return ratę = 0(gKt)P(gxt)) using the median ć estimator provided by MARK to estimate the overdispersion of our data. As the median overdispersion factor (c) was always inferior to 3, we used CJS models for further analyses. However, when median ć was superior to 1, we multiplied the variance-covariance matrix by median Ć to control for the overdispersion of our data.
For both within winter and among year survival analyses, the model with the lowest 2nd order Aikake information criteria (AICc or QAICc when the matrix was multiplied by median 6) was selected as the basie model. In the within winter analyses, the effects of group and period were not significant (see Results 5.5), we therefore used the nuli model (return ratę = <X>(.)p(.)) as basie model for these analyses. Ln the among year analyses, the effect of period was significant on the encounter probability (see Results 5.5), we therefore used the model return ratę = 0(.)P(,) as basie model for the among year analyses.
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